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<front>
<journal-meta>
<journal-id journal-id-type="nlm-ta">PLoS ONE</journal-id>
<journal-id journal-id-type="publisher-id">plos</journal-id>
<journal-id journal-id-type="pmc">plosone</journal-id>
<journal-title-group>
<journal-title>PLOS ONE</journal-title>
</journal-title-group>
<issn pub-type="epub">1932-6203</issn>
<publisher>
<publisher-name>Public Library of Science</publisher-name>
<publisher-loc>San Francisco, CA USA</publisher-loc>
</publisher>
</journal-meta>
<article-meta>
<article-id pub-id-type="doi">10.1371/journal.pone.0180754</article-id>
<article-id pub-id-type="publisher-id">PONE-D-16-45244</article-id>
<article-categories>
<subj-group subj-group-type="heading">
<subject>Research Article</subject>
</subj-group>
<subj-group subj-group-type="Discipline-v3"><subject>Physical sciences</subject><subj-group><subject>Mathematics</subject><subj-group><subject>Applied mathematics</subject><subj-group><subject>Game theory</subject></subj-group></subj-group></subj-group></subj-group><subj-group subj-group-type="Discipline-v3"><subject>Biology and life sciences</subject><subj-group><subject>Psychology</subject><subj-group><subject>Emotions</subject><subj-group><subject>Gratitude</subject></subj-group></subj-group></subj-group></subj-group><subj-group subj-group-type="Discipline-v3"><subject>Social sciences</subject><subj-group><subject>Psychology</subject><subj-group><subject>Emotions</subject><subj-group><subject>Gratitude</subject></subj-group></subj-group></subj-group></subj-group><subj-group subj-group-type="Discipline-v3"><subject>Social sciences</subject><subj-group><subject>Economics</subject><subj-group><subject>Experimental economics</subject></subj-group></subj-group></subj-group><subj-group subj-group-type="Discipline-v3"><subject>Physical sciences</subject><subj-group><subject>Mathematics</subject><subj-group><subject>Applied mathematics</subject><subj-group><subject>Game theory</subject><subj-group><subject>Prisoner's dilemma</subject></subj-group></subj-group></subj-group></subj-group></subj-group><subj-group subj-group-type="Discipline-v3"><subject>Social sciences</subject><subj-group><subject>Sociology</subject><subj-group><subject>Social research</subject></subj-group></subj-group></subj-group><subj-group subj-group-type="Discipline-v3"><subject>Biology and life sciences</subject><subj-group><subject>Behavior</subject></subj-group></subj-group><subj-group subj-group-type="Discipline-v3"><subject>Physical sciences</subject><subj-group><subject>Mathematics</subject><subj-group><subject>Probability theory</subject><subj-group><subject>Markov models</subject></subj-group></subj-group></subj-group></subj-group><subj-group subj-group-type="Discipline-v3"><subject>Biology and life sciences</subject><subj-group><subject>Behavior</subject><subj-group><subject>Recreation</subject><subj-group><subject>Games</subject></subj-group></subj-group></subj-group></subj-group></article-categories>
<title-group>
<article-title>From rationality to cooperativeness: The totally mixed Nash equilibrium in Markov strategies in the iterated Prisoner’s Dilemma</article-title>
<alt-title alt-title-type="running-head">The totally mixed Nash equilibrium in Markov strategies in the iterated Prisoner's Dilemma</alt-title>
</title-group>
<contrib-group>
<contrib contrib-type="author" equal-contrib="yes" xlink:type="simple">
<name name-style="western">
<surname>Menshikov</surname>
<given-names>Ivan S.</given-names>
</name>
<role content-type="http://credit.casrai.org/">Conceptualization</role>
<role content-type="http://credit.casrai.org/">Data curation</role>
<role content-type="http://credit.casrai.org/">Funding acquisition</role>
<role content-type="http://credit.casrai.org/">Methodology</role>
<role content-type="http://credit.casrai.org/">Software</role>
<role content-type="http://credit.casrai.org/">Supervision</role>
<role content-type="http://credit.casrai.org/">Validation</role>
<role content-type="http://credit.casrai.org/">Writing – original draft</role>
<role content-type="http://credit.casrai.org/">Writing – review &amp; editing</role>
<xref ref-type="aff" rid="aff001"><sup>1</sup></xref>
<xref ref-type="aff" rid="aff002"><sup>2</sup></xref>
</contrib>
<contrib contrib-type="author" equal-contrib="yes" xlink:type="simple">
<name name-style="western">
<surname>Shklover</surname>
<given-names>Alexsandr V.</given-names>
</name>
<role content-type="http://credit.casrai.org/">Conceptualization</role>
<role content-type="http://credit.casrai.org/">Formal analysis</role>
<role content-type="http://credit.casrai.org/">Methodology</role>
<role content-type="http://credit.casrai.org/">Software</role>
<role content-type="http://credit.casrai.org/">Supervision</role>
<role content-type="http://credit.casrai.org/">Validation</role>
<xref ref-type="aff" rid="aff001"><sup>1</sup></xref>
</contrib>
<contrib contrib-type="author" corresp="yes" xlink:type="simple">
<contrib-id authenticated="true" contrib-id-type="orcid">http://orcid.org/0000-0002-0504-2643</contrib-id>
<name name-style="western">
<surname>Babkina</surname>
<given-names>Tatiana S.</given-names>
</name>
<role content-type="http://credit.casrai.org/">Data curation</role>
<role content-type="http://credit.casrai.org/">Formal analysis</role>
<role content-type="http://credit.casrai.org/">Investigation</role>
<role content-type="http://credit.casrai.org/">Methodology</role>
<role content-type="http://credit.casrai.org/">Project administration</role>
<role content-type="http://credit.casrai.org/">Resources</role>
<role content-type="http://credit.casrai.org/">Software</role>
<role content-type="http://credit.casrai.org/">Visualization</role>
<role content-type="http://credit.casrai.org/">Writing – original draft</role>
<role content-type="http://credit.casrai.org/">Writing – review &amp; editing</role>
<xref ref-type="aff" rid="aff001"><sup>1</sup></xref>
<xref ref-type="aff" rid="aff003"><sup>3</sup></xref>
<xref ref-type="aff" rid="aff004"><sup>4</sup></xref>
<xref ref-type="fn" rid="econtrib001"><sup>‡</sup></xref>
<xref ref-type="corresp" rid="cor001">*</xref>
</contrib>
<contrib contrib-type="author" xlink:type="simple">
<name name-style="western">
<surname>Myagkov</surname>
<given-names>Mikhail G.</given-names>
</name>
<role content-type="http://credit.casrai.org/">Funding acquisition</role>
<role content-type="http://credit.casrai.org/">Methodology</role>
<role content-type="http://credit.casrai.org/">Project administration</role>
<xref ref-type="aff" rid="aff004"><sup>4</sup></xref>
<xref ref-type="aff" rid="aff005"><sup>5</sup></xref>
<xref ref-type="fn" rid="econtrib001"><sup>‡</sup></xref>
</contrib>
</contrib-group>
<aff id="aff001"><label>1</label> <addr-line>Department of Control and Applied Mathematics, Moscow Institute of Physics and Technology (State University), Moscow, Moscow Region, Russian Federation</addr-line></aff>
<aff id="aff002"><label>2</label> <addr-line>Department of Mathematical Modeling of Economic Systems, Dorodnicyn Computing Center, Federal Research Center «Computer Science and Control» of Russian Academy of Science, Moscow, Moscow Region, Russian Federation</addr-line></aff>
<aff id="aff003"><label>3</label> <addr-line>Center for Design, Manufacturing and Materials, Skolkovo Institute of Science and Technology, Moscow, Moscow Region, Russian Federation</addr-line></aff>
<aff id="aff004"><label>4</label> <addr-line>Laboratory of Experimental Methods in Cognitive and Social Sciences, Tomsk State University, Tomsk, Tomsk Region, Russian Federation</addr-line></aff>
<aff id="aff005"><label>5</label> <addr-line>Department of Political Science, University of Oregon, Eugene, Oregon, United States of America</addr-line></aff>
<contrib-group>
<contrib contrib-type="editor" xlink:type="simple">
<name name-style="western">
<surname>Georgantzis</surname>
<given-names>Nikolaos</given-names>
</name>
<role>Editor</role>
<xref ref-type="aff" rid="edit1"/>
</contrib>
</contrib-group>
<aff id="edit1"><addr-line>Groupe ESC Dijon Bourgogne, FRANCE</addr-line></aff>
<author-notes>
<fn fn-type="conflict" id="coi001">
<p>The authors have declared that no competing interests exist.</p>
</fn>
<fn fn-type="other" id="econtrib001">
<p>‡ These authors also contributed equally to this work.</p>
</fn>
<corresp id="cor001">* E-mail: <email xlink:type="simple">t.babkina@skoltech.ru</email>, <email xlink:type="simple">babkinats@yandex.ru</email></corresp>
</author-notes>
<pub-date pub-type="epub">
<day>30</day>
<month>11</month>
<year>2017</year>
</pub-date>
<pub-date pub-type="collection">
<year>2017</year>
</pub-date>
<volume>12</volume>
<issue>11</issue>
<elocation-id>e0180754</elocation-id>
<history>
<date date-type="received">
<day>14</day>
<month>11</month>
<year>2016</year>
</date>
<date date-type="accepted">
<day>5</day>
<month>10</month>
<year>2017</year>
</date>
</history>
<permissions>
<license xlink:href="https://creativecommons.org/publicdomain/zero/1.0/" xlink:type="simple">
<license-p>This is an open access article, free of all copyright, and may be freely reproduced, distributed, transmitted, modified, built upon, or otherwise used by anyone for any lawful purpose. The work is made available under the <ext-link ext-link-type="uri" xlink:href="https://creativecommons.org/publicdomain/zero/1.0/" xlink:type="simple">Creative Commons CC0</ext-link> public domain dedication.</license-p>
</license>
</permissions>
<self-uri content-type="pdf" xlink:href="info:doi/10.1371/journal.pone.0180754"/>
<abstract>
<p>In this research, the social behavior of the participants in a Prisoner's Dilemma laboratory game is explained on the basis of the quantal response equilibrium concept and the representation of the game in Markov strategies. In previous research, we demonstrated that social interaction during the experiment has a positive influence on cooperation, trust, and gratefulness. This research shows that the quantal response equilibrium concept agrees only with the results of experiments on cooperation in Prisoner’s Dilemma prior to social interaction. However, quantal response equilibrium does not explain of participants’ behavior after social interaction. As an alternative theoretical approach, an examination was conducted of iterated Prisoner's Dilemma game in Markov strategies. We built a totally mixed Nash equilibrium in this game; the equilibrium agrees with the results of the experiments both before and after social interaction.</p>
</abstract>
<funding-group>
<award-group id="award001">
<funding-source>
<institution-wrap>
<institution-id institution-id-type="funder-id">http://dx.doi.org/10.13039/501100002261</institution-id>
<institution>Russian Foundation for Basic Research</institution>
</institution-wrap>
</funding-source>
<award-id>16-01-00633 А</award-id>
<principal-award-recipient>
<name name-style="western">
<surname>Menshikov</surname>
<given-names>Ivan S.</given-names>
</name>
</principal-award-recipient>
</award-group>
<funding-statement>The research was supported by the grant in Russian Foundation for Basic Research, <ext-link ext-link-type="uri" xlink:href="http://www.rfbr.ru/rffi/eng" xlink:type="simple">http://www.rfbr.ru/rffi/eng</ext-link>. The grant number is 16-01-00633 А. The recipient of funding is I.S. Menshikov. Also the conducting of the experiments was supported by Skolkovo Institute of Science and Technology. This research was supported by The Tomsk State University competitiveness improvement program.</funding-statement>
</funding-group>
<counts>
<fig-count count="6"/>
<table-count count="8"/>
<page-count count="17"/>
</counts>
<custom-meta-group>
<custom-meta id="data-availability">
<meta-name>Data Availability</meta-name>
<meta-value>Experimental data are readily available on Harvard Dataverse: <ext-link ext-link-type="uri" xlink:href="https://dx.doi.org/10.7910/DVN/ZGW6ZP" xlink:type="simple">http://dx.doi.org/10.7910/DVN/ZGW6ZP</ext-link>.</meta-value>
</custom-meta>
</custom-meta-group>
</article-meta>
</front>
<body>
<sec id="sec001" sec-type="intro">
<title>Introduction</title>
<p>The traditional approach to analyzing the decision-making process of the participants in game-like interaction is based on the individual rationality principle of each participant [<xref ref-type="bibr" rid="pone.0180754.ref001">1</xref>,<xref ref-type="bibr" rid="pone.0180754.ref002">2</xref>]. The Nash equilibrium and its numerous generalizations postulate the principle of the best response by each participant in the interaction to the behavior of others [<xref ref-type="bibr" rid="pone.0180754.ref003">3</xref>–<xref ref-type="bibr" rid="pone.0180754.ref005">5</xref>].</p>
<p>Such an approach enabled creating and researching numerous models of social and economic behavior noted, in particular, by several Nobel prizewinners in economics. At the same time, extensive empirical and experimental data on game-like interaction have been accumulated. In these data people’s behavior cannot be explained only from the individual rationality position [<xref ref-type="bibr" rid="pone.0180754.ref006">6</xref>–<xref ref-type="bibr" rid="pone.0180754.ref008">8</xref>]. Thus, we must consider the social characteristics of decisions taken:</p>
<list list-type="order">
<list-item><p>cooperation as contrary to individualism [<xref ref-type="bibr" rid="pone.0180754.ref009">9</xref>,<xref ref-type="bibr" rid="pone.0180754.ref010">10</xref>];</p></list-item>
<list-item><p>fairness, based on non-acceptance of inequality [<xref ref-type="bibr" rid="pone.0180754.ref009">9</xref>,<xref ref-type="bibr" rid="pone.0180754.ref011">11</xref>,<xref ref-type="bibr" rid="pone.0180754.ref012">12</xref>];</p></list-item>
<list-item><p>trust and gratefulness [<xref ref-type="bibr" rid="pone.0180754.ref013">13</xref>,<xref ref-type="bibr" rid="pone.0180754.ref014">14</xref>]; and</p></list-item>
<list-item><p>level of social responsibility [<xref ref-type="bibr" rid="pone.0180754.ref015">15</xref>,<xref ref-type="bibr" rid="pone.0180754.ref016">16</xref>].</p></list-item>
</list>
<p>One of the standard methods in the theoretical description of data that does not correspond to the theory of rationality is a quantal response equilibrium (QRE) model. To date there have been several attempts at using the QRE approach in the analysis of experimental data. In [<xref ref-type="bibr" rid="pone.0180754.ref017">17</xref>] it was found that the experimental data on auctions is well interpreted using QRE. Moreover, in [<xref ref-type="bibr" rid="pone.0180754.ref018">18</xref>] it was shown that a QRE model complements the method of maximum likelihood by considering the irrationality of the players participating in experiments. The application of QRE to 2×2 games was researched in [<xref ref-type="bibr" rid="pone.0180754.ref004">4</xref>]. Another approach is the method of Markov chain introduction, demonstrated in [<xref ref-type="bibr" rid="pone.0180754.ref019">19</xref>]. The main problem in such research is the originality of the behavioral experimental data in each piece of research. This demands an individual approach and theoretical basis. In this paper we explain how a QRE model and Markov chains were applied to the data of real experiments with the Prisoner’s Dilemma game and the Trust Game.</p>
</sec>
<sec id="sec002" sec-type="materials|methods">
<title>Materials and methods</title>
<sec id="sec003">
<title>Participants</title>
<p>To analyze the social characteristics of people’s behavior during game-like interaction in small groups (4–12 subjects), numerous experiments were conducted in 2013–2016 at the Laboratory of Experimental Economics (LEE) at the Moscow Institute of Physics and Technology (MIPT) in cooperation with the Skolkovo Institute of Science and Technology in 2013–2016, that clearly reveal one or another social characteristic of behavior. In this paper, the results of eight experiments are presented. In each of them, the number of participants was equal and consisted of 12 people; thus, the data on 96 participants (59 males, 37 females) were taken into consideration. For each experiment, MIPT students who were unknown to each other were selected as participants. Characteristics such as major, group, and year of studies were considered during the selection. Recruitment was by advertisements in the VKontakte social network (<ext-link ext-link-type="uri" xlink:href="http://vk.com" xlink:type="simple">vk.com</ext-link>). Skolkovo Institute of Science and Technology Human Subjects Committee approved the study procedures involving human participants. Written informed consents were obtained from participants. Experimental data are readily available on Harvard Dataverse: <ext-link ext-link-type="uri" xlink:href="https://dx.doi.org/10.7910/DVN/ZGW6ZP" xlink:type="simple">http://dx.doi.org/10.7910/DVN/ZGW6ZP</ext-link>.</p>
</sec>
<sec id="sec004">
<title>Design and procedures</title>
<p>During the experiment the participants were asked to play the following:</p>
<list list-type="order">
<list-item><p>Prisoner’s Dilemma (PD). Each of two participants has two strategies: Cooperation (Up or Left) or Defection (Down or Right). In the standard PD, two players are offered the same points, R, for Cooperation and a smaller gain, P, for Defection. If one of the players cooperates and another defects, the cooperator gains a smaller reward, T, but the defector takes a larger reward, S. Thus, there is a ration between prizes T&gt;R&gt;P&gt;S (<xref ref-type="table" rid="pone.0180754.t001">Table 1</xref>) [<xref ref-type="bibr" rid="pone.0180754.ref006">6</xref>]. Defection is more profitable than Cooperation in any partner’s choice, but mutual Cooperation is more profitable for both than mutual Defection. The Nash equilibrium corresponds to mutual Defection (P, P), but the participants try to establish mutual Cooperation (R, R) [<xref ref-type="bibr" rid="pone.0180754.ref020">20</xref>].</p></list-item>
<list-item><p>Trust and Gratefulness (Trust Game) (TG). One of the participants (the Grantor) can entrust another participant (the Grateful) with some of his or her own money (from 0 to 10). The money obtained (invested) is tripled and the Grateful can share any part of this increased amount with the Grantor (<xref ref-type="fig" rid="pone.0180754.g001">Fig 1</xref>). In the totally mixed Nash equilibrium, there is no sense in gratefulness, and therefore there is no sense in trust, which leads to a zero result for both participants [<xref ref-type="bibr" rid="pone.0180754.ref021">21</xref>,<xref ref-type="bibr" rid="pone.0180754.ref022">22</xref>].</p></list-item>
</list>
<fig id="pone.0180754.g001" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.g001</object-id>
<label>Fig 1</label>
<caption>
<title>The structure of the Trust Game.</title>
<p>The illustration represents the decision making process during the TG. Player 1 starts the game and offers some integer number from zero to ten to player 2. Player 2 gets the offered number multiplied by three. Then player 2 returns any number of the number available.</p>
</caption>
<graphic mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.g001" xlink:type="simple"/>
</fig>
<table-wrap id="pone.0180754.t001" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.t001</object-id>
<label>Table 1</label> <caption><title>Prisoner’s Dilemma payoffs.</title></caption>
<alternatives>
<graphic id="pone.0180754.t001g" mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.t001" xlink:type="simple"/>
<table>
<colgroup>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
</colgroup>
<thead>
<tr>
<th align="center">Payoffs</th>
<th align="center"><italic>Cooperation</italic></th>
<th align="center"><italic>Defection</italic></th>
</tr>
</thead>
<tbody>
<tr style="border-top:thick">
<td align="center"><italic>Cooperation</italic></td>
<td align="center"><bold>R, R</bold></td>
<td align="center"><bold>S, T</bold></td>
</tr>
<tr>
<td align="center"><italic>Defection</italic></td>
<td align="center"><bold>T, S</bold></td>
<td align="center"><bold>P, P</bold></td>
</tr>
</tbody>
</table>
</alternatives>
<table-wrap-foot>
<fn id="t001fn001"><p>For the experiments considered in this paper, the parameters of the PD were set as R = 5, P = 1, S = 0, T = 10 (10 &gt; 5 &gt; 1 &gt; 0).</p></fn>
</table-wrap-foot>
</table-wrap>
<p>In the experiments, gratefulness and trust on average are significantly greater than zero.</p>
<p>Each experiment was divided into three parts:</p>
<p>Part 1, the Anonymous stage. The participants were invited to play 11 rounds in PD at first and then 11 rounds in TG. A specialized tool to design and carry out group experiments in experimental economics, z-Tree developed at the University of Zurich, was used [<xref ref-type="bibr" rid="pone.0180754.ref023">23</xref>]. The participants were able to move to the next round only after all 12 participants made their choices. No one knew who their opponents were and in each round the pairs of participants changed randomly. After each round, the result of the round and the overall result for the current point in the game were displayed on the monitor.</p>
<p>Part 2, the Socialization stage. The participants were invited to take part in interactive cooperation. First the participants memorize each others’ names with the help of a snowball game: they sit in a circle, the first one gives his or her name and a personal characteristic that starts with the same letter as the name, the next participant repeats the name and the characteristic of the first participant and says his or her name and characteristic; then along the chain the game comes to the last person in the circle, who repeats all the names and all the characteristics. Then the participants in reverse order share their personal information: hometown, major, hobby, and interests. Then two captains are chosen as volunteers from among the participants. Other participants must choose the captain whose team they want to join and how many points they are prepared to pay for that. The participants find out their gain for the first part of the game. Then each of the participants except the captains must write on a piece of paper the name of the chosen captain and a number of points from, 0 to 50, that they are ready to pay in order to join the team of the chosen captain. The pieces of paper are personally given to the organizer, who sorts the piece by captains and points. In this way, two teams of four people with captains are formed. The remaining four participants, who paid less than the others, continue as individual participants; they are forbidden to communicate or even look at each other (<xref ref-type="fig" rid="pone.0180754.g002">Fig 2</xref>). The participants are informed about the procedure of distribution by teams beforehand, so all the steps are considered as circumspect and deliberate. At the end of the Socialization stage, the participants in the teams with captains have five minutes to find five common characteristics (eye color, favorite food, movie, etc.) and decide the name of the team.</p>
<fig id="pone.0180754.g002" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.g002</object-id>
<label>Fig 2</label>
<caption>
<title>The illustration of the group formation during the Socialization stage.</title>
<p>Two captains are chosen as volunteers from among the participants. Other participants must choose the captain whose team they want to join and how many points they are prepared to pay for that. The four participants, who paid less than the others, continue as individual participants in Group 3; they are forbidden to communicate or even look at each other. The participants in the teams with captains form Group 1 and Group 2.</p>
</caption>
<graphic mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.g002" xlink:type="simple"/>
</fig>
<p>Part 3, the Socialized stage. After Socialization, the participants are divided into three groups: two groups of four participants with captains and the four participants remaining. In this stage, the participants play PD and TG for 18 rounds within each group, i.e., the participants of group 1 with the captain played only with each other, it was the same for group 2, and the four participants remaining also played only with each other.</p>
<p>Therefore, we have the behavioral data of participants before Socialization in the general group of 12 people and after Socialization in the respective groups of four.</p>
</sec>
<sec id="sec005">
<title>Recent findings</title>
<p>In our study, we focus on the issues connected with the mechanism promoting cooperation. It was shown that the cooperation can be boosted by heterogeneous coupling between interdependent lattices [<xref ref-type="bibr" rid="pone.0180754.ref024">24</xref>], the link weight mechanism [<xref ref-type="bibr" rid="pone.0180754.ref025">25</xref>], and the size of the interaction neighborhood [<xref ref-type="bibr" rid="pone.0180754.ref026">26</xref>]. These findings explain the evolution of cooperation, especially the emergence of cooperation in the non-cooperative games such as PD [<xref ref-type="bibr" rid="pone.0180754.ref027">27</xref>].</p>
<p>Another approach to investigating cooperation is proposed in [<xref ref-type="bibr" rid="pone.0180754.ref001">1</xref>,<xref ref-type="bibr" rid="pone.0180754.ref028">28</xref>–<xref ref-type="bibr" rid="pone.0180754.ref030">30</xref>], where it was shown that incorporating of Socialization in the experiment in PD increases cooperation. The average level of cooperation in Part 1, the Anonymous stage, is 21%, whereas in Part 3, the Socialized stage, the average level of cooperation in the socialized groups is 53% [<xref ref-type="bibr" rid="pone.0180754.ref028">28</xref>]. From the viewpoint of the theory of rationality, as we know, the participants should not choose the strategy of cooperation; therefore the explanation of participants’ behavior in social experiments of this kind does not fit classic economic theory [<xref ref-type="bibr" rid="pone.0180754.ref031">31</xref>]. The increased level of cooperation is explained with the help of incorporating an additional component of the utility function—the social one. In this way, general utility consists of economic (rational) and social utility. The social component is understood as the completion of a socially useful accomplishment. For example, the cooperative move of a participant gives equal gain to an opponent that leads to the increase of social utility. However, it was of interest to us to elicit how the obtained data agrees with other well-known models, so we turned to the idea of QRE.</p>
</sec>
<sec id="sec006">
<title>About quantal response equilibrium</title>
<p>In this section we will discuss the attempts to explain the deviation of the results observed of participants’ behavior from the theoretical Nash equilibrium on the basis of the concept of QRE. We will note that the QRE conception appeared at the intersection of game theory and experimental economics in order to explain behavior of participants in the laboratory experiments that was significantly different from the Nash equilibrium [<xref ref-type="bibr" rid="pone.0180754.ref032">32</xref>,<xref ref-type="bibr" rid="pone.0180754.ref033">33</xref>].</p>
<p>“QRE is an internally consistent equilibrium model, in the sense that the quantal response functions are based on the equilibrium probability distribution of the opponents’ strategy choices rather than simply on arbitrary beliefs the players could have about those probabilities” [<xref ref-type="bibr" rid="pone.0180754.ref032">32</xref>]. One of the model’s features is that it allows game modeling of players who make mistakes. QRE imposes a requirement that beliefs should correspond to the equilibrium choice of probabilities. In this way, QRE demands solutions in the fixated point of choices of probabilities similar to the Nash equilibrium. However, unlike the classic Nash equilibrium, QRE supposes that the pursuit for the best response is realized by participants only in the probabilistic sense: the better the answer is, the higher the probability that it will be chosen by a participant [<xref ref-type="bibr" rid="pone.0180754.ref010">10</xref>,<xref ref-type="bibr" rid="pone.0180754.ref034">34</xref>].</p>
<p>“The QRE has been compared with experimental observation and generally provides a better fit to the data than the NE” [<xref ref-type="bibr" rid="pone.0180754.ref035">35</xref>]. On this basis, we decided to evaluate the model using our experimental data.</p>
<p>According to [<xref ref-type="bibr" rid="pone.0180754.ref033">33</xref>], we introduce QRE through the logistic quantal response function:
<disp-formula id="pone.0180754.e001">
<alternatives>
<graphic id="pone.0180754.e001g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e001" xlink:type="simple"/>
<mml:math display="block" id="M1">
<mml:mrow><mml:msub><mml:mi>σ</mml:mi><mml:mrow><mml:mi>i</mml:mi><mml:mi>j</mml:mi></mml:mrow></mml:msub><mml:mrow><mml:mo>(</mml:mo><mml:mrow><mml:msubsup><mml:mi>u</mml:mi><mml:mi>i</mml:mi><mml:mo>*</mml:mo></mml:msubsup></mml:mrow><mml:mo>)</mml:mo></mml:mrow><mml:mo>=</mml:mo><mml:mfrac><mml:mrow><mml:msup><mml:mi>e</mml:mi><mml:mrow><mml:mi>λ</mml:mi><mml:msub><mml:mi>u</mml:mi><mml:mrow><mml:mi>i</mml:mi><mml:mi>j</mml:mi></mml:mrow></mml:msub></mml:mrow></mml:msup></mml:mrow><mml:mrow><mml:mstyle displaystyle="true"><mml:msubsup><mml:mo>∑</mml:mo><mml:mrow><mml:mi>k</mml:mi><mml:mo>=</mml:mo><mml:mn>1</mml:mn></mml:mrow><mml:mrow><mml:msub><mml:mi>J</mml:mi><mml:mi>i</mml:mi></mml:msub></mml:mrow></mml:msubsup><mml:mrow><mml:msup><mml:mi>e</mml:mi><mml:mrow><mml:mi>λ</mml:mi><mml:msub><mml:mi>u</mml:mi><mml:mrow><mml:mi>i</mml:mi><mml:mi>k</mml:mi></mml:mrow></mml:msub></mml:mrow></mml:msup></mml:mrow></mml:mstyle></mml:mrow></mml:mfrac></mml:mrow>
</mml:math>
</alternatives>
<label>(1)</label>
</disp-formula></p>
<p>Here u<sub>ij</sub> is the expected payoff of player i with strategy j, (j∈{1,…,J<sub>i</sub>}), <inline-formula id="pone.0180754.e002"><alternatives><graphic id="pone.0180754.e002g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e002" xlink:type="simple"/><mml:math display="inline" id="M2"><mml:mrow><mml:msubsup><mml:mi>u</mml:mi><mml:mi>i</mml:mi><mml:mo>*</mml:mo></mml:msubsup><mml:mo>=</mml:mo><mml:mo stretchy="false">(</mml:mo><mml:msub><mml:mi>u</mml:mi><mml:mrow><mml:mi>i</mml:mi><mml:mn>1</mml:mn></mml:mrow></mml:msub><mml:mo>,</mml:mo><mml:mo>…</mml:mo><mml:mo>,</mml:mo><mml:msub><mml:mi>u</mml:mi><mml:mrow><mml:mi>i</mml:mi><mml:msub><mml:mi>J</mml:mi><mml:mi>i</mml:mi></mml:msub></mml:mrow></mml:msub><mml:mo stretchy="false">)</mml:mo></mml:mrow></mml:math></alternatives></inline-formula> “If each player uses a logistic response function, QRE or logit equilibria are the solutions of Π<sub>ij</sub> = α<sub>ij</sub>, where Π<sub>ij</sub> is the frequency of strategy j in player i” [<xref ref-type="bibr" rid="pone.0180754.ref036">36</xref>].</p>
</sec>
<sec id="sec007">
<title>QRE in the PD game</title>
<p>For the PD game that we considered, the QRE (<xref ref-type="table" rid="pone.0180754.t001">Table 1</xref>) could be determined as follows. Let p be the probability of the cooperative move of a partner, then the expected gain from the cooperative action equals 5p+0(1-p) = 5p and the expected gain from uncooperative action equals 10p+1(1-p) = 9p+1 [<xref ref-type="bibr" rid="pone.0180754.ref004">4</xref>]. We define as the precision parameter, which is inversely related to the variance of the error (2). For every λ, p = QRE(λ) could be found from the formula (<xref ref-type="disp-formula" rid="pone.0180754.e001">1</xref>) from the solution of the Eq (<xref ref-type="disp-formula" rid="pone.0180754.e003">2</xref>)
<disp-formula id="pone.0180754.e003">
<alternatives>
<graphic id="pone.0180754.e003g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e003" xlink:type="simple"/>
<mml:math display="block" id="M3">
<mml:mrow><mml:mfrac><mml:mrow><mml:mi mathvariant="normal">exp</mml:mi><mml:mrow><mml:mo>(</mml:mo><mml:mrow><mml:mi>λ</mml:mi><mml:mo>×</mml:mo><mml:mn>5</mml:mn><mml:mi>p</mml:mi></mml:mrow><mml:mo>)</mml:mo></mml:mrow></mml:mrow><mml:mrow><mml:mi mathvariant="normal">exp</mml:mi><mml:mrow><mml:mo>(</mml:mo><mml:mrow><mml:mi>λ</mml:mi><mml:mo>×</mml:mo><mml:mn>5</mml:mn><mml:mi>p</mml:mi></mml:mrow><mml:mo>)</mml:mo></mml:mrow><mml:mo>+</mml:mo><mml:mi mathvariant="normal">exp</mml:mi><mml:mrow><mml:mo>(</mml:mo><mml:mrow><mml:mi>λ</mml:mi><mml:mrow><mml:mo>(</mml:mo><mml:mrow><mml:mn>9</mml:mn><mml:mi>p</mml:mi><mml:mo>+</mml:mo><mml:mn>1</mml:mn></mml:mrow><mml:mo>)</mml:mo></mml:mrow></mml:mrow><mml:mo>)</mml:mo></mml:mrow></mml:mrow></mml:mfrac><mml:mo>=</mml:mo><mml:mfrac><mml:mn>1</mml:mn><mml:mrow><mml:mn>1</mml:mn><mml:mo>+</mml:mo><mml:mi mathvariant="normal">exp</mml:mi><mml:mrow><mml:mo>(</mml:mo><mml:mrow><mml:mi>λ</mml:mi><mml:mrow><mml:mo>(</mml:mo><mml:mrow><mml:mn>4</mml:mn><mml:mi>p</mml:mi><mml:mo>+</mml:mo><mml:mn>1</mml:mn></mml:mrow><mml:mo>)</mml:mo></mml:mrow></mml:mrow><mml:mo>)</mml:mo></mml:mrow></mml:mrow></mml:mfrac></mml:mrow>
</mml:math>
</alternatives>
<label>(2)</label>
</disp-formula></p>
<p>At λ = 0 the probability of the cooperative move by QRE(λ) equals 0.5 (the chaotic behavior). With an increase of λthe probability of cooperation by QRE(λ) decreases and within the limit λ→∞ strives to 0 and this corresponds to the single Nash equilibrium in PD. Thus, from the QRE position any percentage of cooperative moves less than 50% could be justified. It is quite suitable for games before Socialization. Mathematically that means the solution of the QRE(λ) = p equation is relative to the λ parameter for a given observed level of the cooperative moves p. In this case, this equation is easily solved:
<disp-formula id="pone.0180754.e004">
<alternatives>
<graphic id="pone.0180754.e004g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e004" xlink:type="simple"/>
<mml:math display="block" id="M4">
<mml:mrow><mml:mi>λ</mml:mi><mml:mo>=</mml:mo><mml:mfrac><mml:mrow><mml:mi mathvariant="normal">ln</mml:mi><mml:mo stretchy="false">(</mml:mo><mml:mfrac><mml:mrow><mml:mn>1</mml:mn><mml:mo>−</mml:mo><mml:mi>p</mml:mi></mml:mrow><mml:mi>p</mml:mi></mml:mfrac><mml:mo stretchy="false">)</mml:mo></mml:mrow><mml:mrow><mml:mn>4</mml:mn><mml:mi>p</mml:mi><mml:mo>+</mml:mo><mml:mn>1</mml:mn></mml:mrow></mml:mfrac></mml:mrow>
</mml:math>
</alternatives>
<label>(3)</label>
</disp-formula></p>
<p>We give all the solutions to this equation for the series of experiments (<xref ref-type="table" rid="pone.0180754.t002">Table 2</xref>) in fall, 2015.</p>
<table-wrap id="pone.0180754.t002" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.t002</object-id>
<label>Table 2</label> <caption><title>The average level of cooperation and the QRE parameter for all the experiments in PD before the Socialization stage.</title></caption>
<alternatives>
<graphic id="pone.0180754.t002g" mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.t002" xlink:type="simple"/>
<table>
<colgroup>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
</colgroup>
<thead>
<tr>
<th align="center" rowspan="2">Experiment date</th>
<th align="center" colspan="2">The average level of cooperation before Socialization (%)</th>
<th align="center" rowspan="2">Lambda, λ</th>
</tr>
<tr>
<th align="center">M</th>
<th align="center">SD</th>
</tr>
</thead>
<tbody>
<tr style="border-top:thick">
<td align="center">15.09.2015</td>
<td align="center">0.121</td>
<td align="center">0.13</td>
<td align="center">1.334</td>
</tr>
<tr>
<td align="center">21.09.2015</td>
<td align="center">0.144</td>
<td align="center">0.15</td>
<td align="center">1.131</td>
</tr>
<tr>
<td align="center">28.09.2015</td>
<td align="center">0.409</td>
<td align="center">0.31</td>
<td align="center">0.139</td>
</tr>
<tr>
<td align="center">05.10.2015</td>
<td align="center">0.227</td>
<td align="center">0.14</td>
<td align="center">0.641</td>
</tr>
<tr>
<td align="center">09.10.2015</td>
<td align="center">0.364</td>
<td align="center">0.21</td>
<td align="center">0.228</td>
</tr>
<tr>
<td align="center">12.10.2015</td>
<td align="center">0.417</td>
<td align="center">0.26</td>
<td align="center">0.126</td>
</tr>
<tr>
<td align="center">19.10.2015</td>
<td align="center">0.356</td>
<td align="center">0.26</td>
<td align="center">0.244</td>
</tr>
<tr>
<td align="center">26.10.2015</td>
<td align="center">0.212</td>
<td align="center">0.22</td>
<td align="center">0.710</td>
</tr>
</tbody>
</table>
</alternatives>
<table-wrap-foot>
<fn id="t002fn001"><p>The table represents the average level of cooperation before Socialization and calculated Lambda for all experiments.</p></fn>
</table-wrap-foot>
</table-wrap>
<p>For clarity, we represent this dependence graphically (<xref ref-type="fig" rid="pone.0180754.g003">Fig 3</xref>).</p>
<fig id="pone.0180754.g003" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.g003</object-id>
<label>Fig 3</label>
<caption>
<title>The average level of cooperation and the QRE parameter for all experiments in PD before the Socialization stage.</title>
<p>The λparameter (y-axis) as a function of the average level of cooperation in every experiments (x-axis).</p>
</caption>
<graphic mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.g003" xlink:type="simple"/>
</fig>
<p>We see that the maximum degree of cooperation before Socialization was achieved 12.10.2015 and is 41.7%, which corresponds to the significantly positive value λ = 0.126. The minimum degree of cooperation before Socialization was reached 15.09.2015 and is 12.1% at λ = 1.334 (<xref ref-type="fig" rid="pone.0180754.g003">Fig 3</xref>), which is far from the limit value. The average value of cooperation in the experiment series is 28% at λ = 0.44.</p>
<p>Thus, the calculations show that the behavior of the participants of the experiments in PD before Socialization is completely described by the QRE concept, which is the adopted deviation from the Nash equilibrium towards the easing of the requirements of the best answer.</p>
<p>However, after Socialization the situation radically changes.</p>
<p>As <xref ref-type="table" rid="pone.0180754.t003">Table 3</xref> shows, the level of cooperation after Socialization is over 50% in all the experiments, which is why the QRE concept is not fully applicable in this case.</p>
<table-wrap id="pone.0180754.t003" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.t003</object-id>
<label>Table 3</label> <caption><title>The average levels of the cooperative moves in PD after Socialization.</title></caption>
<alternatives>
<graphic id="pone.0180754.t003g" mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.t003" xlink:type="simple"/>
<table>
<colgroup>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
</colgroup>
<thead>
<tr>
<th align="center" rowspan="2">Experiment date</th>
<th align="center" colspan="2">The average level of cooperation after Socialization (%)</th>
</tr>
<tr>
<th align="center">M</th>
<th align="center">SD</th>
</tr>
</thead>
<tbody>
<tr style="border-top:thick">
<td align="center">15.09.2015</td>
<td align="center">0.741</td>
<td align="center">0.35</td>
</tr>
<tr>
<td align="center">21.09.2015</td>
<td align="center">0.537</td>
<td align="center">0.33</td>
</tr>
<tr>
<td align="center">28.09.2015</td>
<td align="center">0.639</td>
<td align="center">0.43</td>
</tr>
<tr>
<td align="center">05.10.2015</td>
<td align="center">0.574</td>
<td align="center">0.35</td>
</tr>
<tr>
<td align="center">09.10.2015</td>
<td align="center">0.528</td>
<td align="center">0.29</td>
</tr>
<tr>
<td align="center">12.10.2015</td>
<td align="center">0.718</td>
<td align="center">0.33</td>
</tr>
<tr>
<td align="center">19.10.2015</td>
<td align="center">0.796</td>
<td align="center">0.27</td>
</tr>
<tr>
<td align="center">26.10.2015</td>
<td align="center">0.556</td>
<td align="center">0.37</td>
</tr>
</tbody>
</table>
</alternatives>
<table-wrap-foot>
<fn id="t003fn001"><p>The table represents the average level of cooperation after Socialization.</p></fn>
</table-wrap-foot>
</table-wrap>
<p>This means that it is necessary to search for an alternative theoretical game model for the behavior of participants.</p>
</sec>
<sec id="sec008">
<title>QRE in the TG</title>
<p>Let us find the QRE for the TG, which was also in a series of experiments in fall, 2015.</p>
<p>Unlike the static PD game, TG is a dynamic game with perfect information. The QRE concept is theoretically applicable in this case too. Let k = 0,…,10 be the trust level of player 1, and n = 0,…,3k be the gratefulness level of player 2. For a given level of k,n the winning of player 1 is 10-n+k, and the winning of player 2 is 3k-n. According to QRE, the probability p<sub>2</sub>(k,n)of thanks of a level n for a given level of trust k is determined by the formula
<disp-formula id="pone.0180754.e005">
<alternatives>
<graphic id="pone.0180754.e005g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e005" xlink:type="simple"/>
<mml:math display="block" id="M5">
<mml:mrow><mml:msub><mml:mi>p</mml:mi><mml:mn>2</mml:mn></mml:msub><mml:mo stretchy="false">(</mml:mo><mml:mi>k</mml:mi><mml:mo>,</mml:mo><mml:mi>n</mml:mi><mml:mo stretchy="false">)</mml:mo><mml:mo>=</mml:mo><mml:mfrac><mml:mrow><mml:msup><mml:mi>e</mml:mi><mml:mrow><mml:mi>λ</mml:mi><mml:mo stretchy="false">(</mml:mo><mml:mn>3</mml:mn><mml:mi>k</mml:mi><mml:mo>−</mml:mo><mml:mi>n</mml:mi><mml:mo stretchy="false">)</mml:mo></mml:mrow></mml:msup></mml:mrow><mml:mrow><mml:mstyle displaystyle="true"><mml:munderover><mml:mo>∑</mml:mo><mml:mrow><mml:mi>i</mml:mi><mml:mo>=</mml:mo><mml:mn>0</mml:mn></mml:mrow><mml:mrow><mml:mn>3</mml:mn><mml:mi>k</mml:mi></mml:mrow></mml:munderover><mml:mrow><mml:msup><mml:mi>e</mml:mi><mml:mrow><mml:mi>λ</mml:mi><mml:mo stretchy="false">(</mml:mo><mml:mn>3</mml:mn><mml:mi>k</mml:mi><mml:mo>−</mml:mo><mml:mi>i</mml:mi><mml:mo stretchy="false">)</mml:mo></mml:mrow></mml:msup></mml:mrow></mml:mstyle></mml:mrow></mml:mfrac></mml:mrow>
</mml:math>
</alternatives>
<label>(4)</label>
</disp-formula></p>
<p>For this reason, the expected winning u<sub>1</sub>(k)of player 1 with a level of trust k is
<disp-formula id="pone.0180754.e006">
<alternatives>
<graphic id="pone.0180754.e006g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e006" xlink:type="simple"/>
<mml:math display="block" id="M6">
<mml:mrow><mml:msub><mml:mi>u</mml:mi><mml:mn>1</mml:mn></mml:msub><mml:mo stretchy="false">(</mml:mo><mml:mi>k</mml:mi><mml:mo stretchy="false">)</mml:mo><mml:mo>=</mml:mo><mml:mn>10</mml:mn><mml:mo>−</mml:mo><mml:mi>k</mml:mi><mml:mo>+</mml:mo><mml:mstyle displaystyle="true"><mml:munderover><mml:mo>∑</mml:mo><mml:mrow><mml:mi>n</mml:mi><mml:mo>=</mml:mo><mml:mn>0</mml:mn></mml:mrow><mml:mrow><mml:mn>3</mml:mn><mml:mi>k</mml:mi></mml:mrow></mml:munderover><mml:mrow><mml:mi>n</mml:mi><mml:msub><mml:mi>p</mml:mi><mml:mn>2</mml:mn></mml:msub><mml:mo stretchy="false">(</mml:mo><mml:mi>k</mml:mi><mml:mo>,</mml:mo><mml:mi>n</mml:mi><mml:mo stretchy="false">)</mml:mo></mml:mrow></mml:mstyle></mml:mrow>
</mml:math>
</alternatives>
<label>(5)</label>
</disp-formula></p>
<p>Then the probability p<sub>1</sub>(k)of trust of level k in QRE is determined by the formula
<disp-formula id="pone.0180754.e007">
<alternatives>
<graphic id="pone.0180754.e007g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e007" xlink:type="simple"/>
<mml:math display="block" id="M7">
<mml:mrow><mml:msub><mml:mi>p</mml:mi><mml:mn>1</mml:mn></mml:msub><mml:mo stretchy="false">(</mml:mo><mml:mi>k</mml:mi><mml:mo stretchy="false">)</mml:mo><mml:mo>=</mml:mo><mml:mfrac><mml:mrow><mml:msup><mml:mi>e</mml:mi><mml:mrow><mml:mi>λ</mml:mi><mml:msub><mml:mi>u</mml:mi><mml:mn>1</mml:mn></mml:msub><mml:mo stretchy="false">(</mml:mo><mml:mi>k</mml:mi><mml:mo stretchy="false">)</mml:mo></mml:mrow></mml:msup></mml:mrow><mml:mrow><mml:mstyle displaystyle="true"><mml:munderover><mml:mo>∑</mml:mo><mml:mrow><mml:mi>i</mml:mi><mml:mo>=</mml:mo><mml:mn>0</mml:mn></mml:mrow><mml:mrow><mml:mn>10</mml:mn></mml:mrow></mml:munderover><mml:mrow><mml:msup><mml:mi>e</mml:mi><mml:mrow><mml:mi>λ</mml:mi><mml:msub><mml:mi>u</mml:mi><mml:mn>1</mml:mn></mml:msub><mml:mo stretchy="false">(</mml:mo><mml:mi>i</mml:mi><mml:mo stretchy="false">)</mml:mo></mml:mrow></mml:msup></mml:mrow></mml:mstyle></mml:mrow></mml:mfrac></mml:mrow>
</mml:math>
</alternatives>
<label>(6)</label>
</disp-formula></p>
<p>To find the parameter λ according to the results of the experiment let us calculate the average levels of trust k* and thanks n* and compare them with the theoretical expected level of trust k(λ) and gratefulness n(λ), which are calculated as
<disp-formula id="pone.0180754.e008">
<alternatives>
<graphic id="pone.0180754.e008g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e008" xlink:type="simple"/>
<mml:math display="block" id="M8">
<mml:mrow><mml:mi>k</mml:mi><mml:mo stretchy="false">(</mml:mo><mml:mi>λ</mml:mi><mml:mo stretchy="false">)</mml:mo><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:munderover><mml:mo>∑</mml:mo><mml:mrow><mml:mi>k</mml:mi><mml:mo>=</mml:mo><mml:mn>0</mml:mn></mml:mrow><mml:mrow><mml:mn>10</mml:mn></mml:mrow></mml:munderover><mml:mrow><mml:msub><mml:mi>p</mml:mi><mml:mn>1</mml:mn></mml:msub><mml:mo stretchy="false">(</mml:mo><mml:mi>k</mml:mi><mml:mo stretchy="false">)</mml:mo><mml:mo>⋅</mml:mo><mml:mi>k</mml:mi></mml:mrow></mml:mstyle><mml:mo>,</mml:mo><mml:mspace width="0.5em"/><mml:mi>n</mml:mi><mml:mo stretchy="false">(</mml:mo><mml:mi>λ</mml:mi><mml:mo stretchy="false">)</mml:mo><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:munderover><mml:mo>∑</mml:mo><mml:mrow><mml:mi>k</mml:mi><mml:mo>=</mml:mo><mml:mn>0</mml:mn></mml:mrow><mml:mrow><mml:mn>10</mml:mn></mml:mrow></mml:munderover><mml:mrow><mml:mstyle displaystyle="true"><mml:munderover><mml:mo>∑</mml:mo><mml:mrow><mml:mi>n</mml:mi><mml:mo>=</mml:mo><mml:mn>0</mml:mn></mml:mrow><mml:mrow><mml:mn>3</mml:mn><mml:mi>k</mml:mi></mml:mrow></mml:munderover><mml:mrow><mml:msub><mml:mi>p</mml:mi><mml:mn>1</mml:mn></mml:msub><mml:mo stretchy="false">(</mml:mo><mml:mi>k</mml:mi><mml:mo stretchy="false">)</mml:mo><mml:mo>⋅</mml:mo><mml:msub><mml:mi>p</mml:mi><mml:mn>2</mml:mn></mml:msub><mml:mo stretchy="false">(</mml:mo><mml:mi>k</mml:mi><mml:mo>,</mml:mo><mml:mi>n</mml:mi><mml:mo stretchy="false">)</mml:mo><mml:mo>⋅</mml:mo><mml:mi>n</mml:mi></mml:mrow></mml:mstyle></mml:mrow></mml:mstyle></mml:mrow>
</mml:math>
</alternatives>
<label>(7)</label>
</disp-formula></p>
<p>Let us select parameter λ so that several levels (k(λ),n(λ)) would become as close as possible to the levels (k*,n*) observed in the experiment.</p>
<p>As the following results of calculations show, parameter λ even for games before Socialization is rather close to 0; thus the behavior of the participants according to QRE is treated as nearly chaotic for some experiments.</p>
<p><xref ref-type="table" rid="pone.0180754.t004">Table 4</xref> shows the average values of trust and gratefulness for each experiment in the fall, 2015 series with the calculated parameter λ and approximate values of trust and gratefulness, which approximate the specified average values in the best way. The average of value λ for this series is estimated at 0.17, which is significantly lower than the average value λ obtained previously for the PD game in the same series of experiments. Hence, we can conclude that the QRE concept poorly explains the results of experiments even before Socialization.</p>
<table-wrap id="pone.0180754.t004" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.t004</object-id>
<label>Table 4</label> <caption><title>Average trust and gratefulness in comparison with QRE for the TG.</title></caption>
<alternatives>
<graphic id="pone.0180754.t004g" mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.t004" xlink:type="simple"/>
<table>
<colgroup>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
</colgroup>
<thead>
<tr>
<th align="center" rowspan="2">Experiment date</th>
<th align="center" colspan="2">Average trust</th>
<th align="center" colspan="2">Average gratefulness</th>
<th align="center" rowspan="2">Lambda</th>
<th align="center" rowspan="2">Forecast of trust</th>
<th align="center" rowspan="2">Forecast of gratefulness</th>
</tr>
<tr>
<th align="center">M</th>
<th align="center">SD</th>
<th align="center">M</th>
<th align="center">SD</th>
</tr>
</thead>
<tbody>
<tr style="border-top:thick">
<td align="center">15.09.2015</td>
<td align="center">5.38</td>
<td align="center">3.48</td>
<td align="center">3.17</td>
<td align="center">2.54</td>
<td align="center">0.15</td>
<td align="center">4.39</td>
<td align="center">3.63</td>
</tr>
<tr>
<td align="center">21.09.2015</td>
<td align="center">4.25</td>
<td align="center">2.68</td>
<td align="center">1.98</td>
<td align="center">2.15</td>
<td align="center">0.23</td>
<td align="center">3.67</td>
<td align="center">2.43</td>
</tr>
<tr>
<td align="center">05.10.2015</td>
<td align="center">3.33</td>
<td align="center">1.36</td>
<td align="center">0.94</td>
<td align="center">0.69</td>
<td align="center">0.34</td>
<td align="center">2.81</td>
<td align="center">1.55</td>
</tr>
<tr>
<td align="center">09.10.2015</td>
<td align="center">5.58</td>
<td align="center">2.74</td>
<td align="center">4.12</td>
<td align="center">3.27</td>
<td align="center">0.11</td>
<td align="center">4.70</td>
<td align="center">4.40</td>
</tr>
<tr>
<td align="center">12.10.2015</td>
<td align="center">5.80</td>
<td align="center">3.03</td>
<td align="center">4.87</td>
<td align="center">3.75</td>
<td align="center">0.09</td>
<td align="center">4.89</td>
<td align="center">5.08</td>
</tr>
<tr>
<td align="center">19.10.2015</td>
<td align="center">5.81</td>
<td align="center">3.32</td>
<td align="center">6.88</td>
<td align="center">3.29</td>
<td align="center">0.03</td>
<td align="center">5.08</td>
<td align="center">6.86</td>
</tr>
<tr>
<td align="center">26.10.2015</td>
<td align="center">4.06</td>
<td align="center">3.28</td>
<td align="center">2.62</td>
<td align="center">2.07</td>
<td align="center">0.21</td>
<td align="center">3.89</td>
<td align="center">2.74</td>
</tr>
</tbody>
</table>
</alternatives>
<table-wrap-foot>
<fn id="t004fn001"><p>The table shows the average trust, average gratefulness, calculated Lambda, forecast of trust and forecast of gratefulness for all experiments.</p></fn>
</table-wrap-foot>
</table-wrap>
</sec>
<sec id="sec009">
<title>Model of iterated PD in Markov strategies</title>
<p>Let us construct and analyze a model of the iterated PD in Markov strategies. At the beginning, take into consideration the effect of the iterated PD several times with random partners. For simplicity let us assume that every participant responds only to the move made by his or her partner in the previous round. Such strategies are called Markov strategies or strategies with memory length equal to one [<xref ref-type="bibr" rid="pone.0180754.ref037">37</xref>,<xref ref-type="bibr" rid="pone.0180754.ref038">38</xref>]. Previously, the Markov chains were used more than once to find the equilibria for the Prisoner's Dilemma [<xref ref-type="bibr" rid="pone.0180754.ref019">19</xref>,<xref ref-type="bibr" rid="pone.0180754.ref037">37</xref>–<xref ref-type="bibr" rid="pone.0180754.ref040">40</xref>]. In [<xref ref-type="bibr" rid="pone.0180754.ref019">19</xref>] only equilibria for "good", cooperative strategies were found. Based on the past results, we also decided to apply Markov strategies for the theoretical justification of the experimental data. However, we were interested not in extreme cases, which lead to total cooperation rarely observed in experiments, but in the internal equilibrium points, in which both cooperation and betrayal are selected with positive probabilities.</p>
<p>For the PD game after Socialization, the following approach described in previous works [<xref ref-type="bibr" rid="pone.0180754.ref038">38</xref>,<xref ref-type="bibr" rid="pone.0180754.ref041">41</xref>–<xref ref-type="bibr" rid="pone.0180754.ref045">45</xref>] was suitable the most:</p>
<p>Let γ<sub>i</sub> denote reciprocal cooperation, i.e. the probability that a participant i will act cooperatively after the previous round in which his or her partner played cooperatively.</p>
<p>Let α<sub>i</sub> denote tolerance to defection, i.e. the probability that a participant i will act cooperatively after the previous round in which his or her partner played non-cooperatively.</p>
<p>For the given parameters of cooperation and tolerance of the pair of participants, we obtain a Markov process with a finite number of states [<xref ref-type="bibr" rid="pone.0180754.ref046">46</xref>–<xref ref-type="bibr" rid="pone.0180754.ref048">48</xref>]. In the stationary distribution, each player of the pair of participants will be in one of two possible states: {Cooperation, Defection}. The stationary probability p<sub>i</sub><sup>c</sup> for a participant i to be in a cooperative state depends on the stationary probability p<sub>j</sub><sup>c</sup> of a participant j≠i and strategic parameters of reciprocal cooperation γ<sub>i</sub> and tolerance to defection α<sub>i</sub> of a participant i in the following way
<disp-formula id="pone.0180754.e009">
<alternatives>
<graphic id="pone.0180754.e009g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e009" xlink:type="simple"/>
<mml:math display="block" id="M9">
<mml:mrow><mml:msubsup><mml:mi>p</mml:mi><mml:mi>i</mml:mi><mml:mi>c</mml:mi></mml:msubsup><mml:mo>=</mml:mo><mml:msub><mml:mi>γ</mml:mi><mml:mi>i</mml:mi></mml:msub><mml:msubsup><mml:mi>p</mml:mi><mml:mi>j</mml:mi><mml:mi>c</mml:mi></mml:msubsup><mml:mo>+</mml:mo><mml:msub><mml:mi>α</mml:mi><mml:mi>i</mml:mi></mml:msub><mml:mo stretchy="false">(</mml:mo><mml:mn>1</mml:mn><mml:mo>−</mml:mo><mml:msubsup><mml:mi>p</mml:mi><mml:mi>j</mml:mi><mml:mi>c</mml:mi></mml:msubsup><mml:mo stretchy="false">)</mml:mo><mml:mo>,</mml:mo><mml:mspace width="0.5em"/><mml:mi>i</mml:mi><mml:mo>,</mml:mo><mml:mi>j</mml:mi><mml:mo>=</mml:mo><mml:mn>1</mml:mn><mml:mo>,</mml:mo><mml:mn>2</mml:mn><mml:mo>,</mml:mo><mml:mspace width="0.25em"/><mml:mi>j</mml:mi><mml:mo>≠</mml:mo><mml:mi>i</mml:mi></mml:mrow>
</mml:math>
</alternatives>
<label>(8)</label>
</disp-formula></p>
<p>For the given strategies {α<sub>1</sub>, α<sub>2</sub>, γ<sub>1,</sub> γ<sub>2</sub>} of two unknowns participants (1 –the first participant and 2 –the second) this system of two linear equations with is easily solved in explicit form:
<disp-formula id="pone.0180754.e010">
<alternatives>
<graphic id="pone.0180754.e010g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e010" xlink:type="simple"/>
<mml:math display="block" id="M10">
<mml:mrow><mml:msubsup><mml:mi>p</mml:mi><mml:mi>i</mml:mi><mml:mi>c</mml:mi></mml:msubsup><mml:mo>=</mml:mo><mml:mfrac><mml:mrow><mml:msub><mml:mi>α</mml:mi><mml:mi>i</mml:mi></mml:msub><mml:mo>−</mml:mo><mml:msub><mml:mi>α</mml:mi><mml:mi>j</mml:mi></mml:msub><mml:mo stretchy="false">(</mml:mo><mml:msub><mml:mi>α</mml:mi><mml:mi>i</mml:mi></mml:msub><mml:mo>−</mml:mo><mml:msub><mml:mi>γ</mml:mi><mml:mi>i</mml:mi></mml:msub><mml:mo stretchy="false">)</mml:mo></mml:mrow><mml:mrow><mml:mn>1</mml:mn><mml:mo>−</mml:mo><mml:mo stretchy="false">(</mml:mo><mml:msub><mml:mi>α</mml:mi><mml:mn>1</mml:mn></mml:msub><mml:mo>−</mml:mo><mml:msub><mml:mi>γ</mml:mi><mml:mn>1</mml:mn></mml:msub><mml:mo stretchy="false">)</mml:mo><mml:mo stretchy="false">(</mml:mo><mml:msub><mml:mi>α</mml:mi><mml:mn>2</mml:mn></mml:msub><mml:mo>−</mml:mo><mml:msub><mml:mi>γ</mml:mi><mml:mn>2</mml:mn></mml:msub><mml:mo stretchy="false">)</mml:mo></mml:mrow></mml:mfrac><mml:mo>,</mml:mo><mml:mspace width="0.25em"/><mml:mi>i</mml:mi><mml:mo>=</mml:mo><mml:mn>1</mml:mn><mml:mo>,</mml:mo><mml:mn>2</mml:mn></mml:mrow>
</mml:math>
</alternatives>
<label>(9)</label>
</disp-formula></p>
<p>The composition of corresponding probabilities gives the stationary distribution for all four pairs of actions of the participants, and based on this distribution we can calculate the profits of the participants. Omitting the intermediate calculations, let us write the expression for the winning of participant 1:
<disp-formula id="pone.0180754.e011">
<alternatives>
<graphic id="pone.0180754.e011g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e011" xlink:type="simple"/>
<mml:math display="block" id="M11">
<mml:mrow><mml:msub><mml:mi>U</mml:mi><mml:mn>1</mml:mn></mml:msub><mml:mo stretchy="false">(</mml:mo><mml:msubsup><mml:mi>p</mml:mi><mml:mn>1</mml:mn><mml:mi>c</mml:mi></mml:msubsup><mml:mo>,</mml:mo><mml:msubsup><mml:mi>p</mml:mi><mml:mn>2</mml:mn><mml:mi>c</mml:mi></mml:msubsup><mml:mo stretchy="false">)</mml:mo><mml:mo>=</mml:mo><mml:mo>−</mml:mo><mml:mn>4</mml:mn><mml:msubsup><mml:mi>p</mml:mi><mml:mn>1</mml:mn><mml:mi>c</mml:mi></mml:msubsup><mml:msubsup><mml:mi>p</mml:mi><mml:mn>2</mml:mn><mml:mi>c</mml:mi></mml:msubsup><mml:mo>−</mml:mo><mml:msubsup><mml:mi>p</mml:mi><mml:mn>1</mml:mn><mml:mi>c</mml:mi></mml:msubsup><mml:mo>+</mml:mo><mml:mn>9</mml:mn><mml:msubsup><mml:mi>p</mml:mi><mml:mn>2</mml:mn><mml:mi>c</mml:mi></mml:msubsup><mml:mo>+</mml:mo><mml:mn>1</mml:mn></mml:mrow>
</mml:math>
</alternatives>
<label>(10)</label>
</disp-formula></p>
<p>It should be remembered that (p<sub>1</sub><sup>c</sup>, p<sub>2</sub><sup>c</sup>) in their turn depend on {α<sub>1</sub>,α<sub>2</sub>,γ<sub>1</sub>,γ<sub>2</sub>} as indicated above. Therefore, there turns out to be some kind of a game in a normal form with nonlinear payoff functions. But a symmetric totally mixed Nash equilibria (in Markov strategies) {α, α, γ, γ} can be found in explicit form in this game:
<disp-formula id="pone.0180754.e012">
<alternatives>
<graphic id="pone.0180754.e012g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e012" xlink:type="simple"/>
<mml:math display="block" id="M12">
<mml:mrow><mml:mn>5</mml:mn><mml:msup><mml:mi>α</mml:mi><mml:mn>2</mml:mn></mml:msup><mml:mo>+</mml:mo><mml:mn>9</mml:mn><mml:msup><mml:mi>γ</mml:mi><mml:mn>2</mml:mn></mml:msup><mml:mo>−</mml:mo><mml:mn>14</mml:mn><mml:mi>α</mml:mi><mml:mi>γ</mml:mi><mml:mo>+</mml:mo><mml:mn>14</mml:mn><mml:mi>α</mml:mi><mml:mo>−</mml:mo><mml:mn>10</mml:mn><mml:mi>γ</mml:mi><mml:mo>+</mml:mo><mml:mn>1</mml:mn><mml:mo>=</mml:mo><mml:mn>0</mml:mn></mml:mrow>
</mml:math>
</alternatives>
<label>(11)</label>
</disp-formula></p>
<p>It can be checked whether this curve of the second order is a hyperbolic curve. Let us represent it in the intersection with a single square of tolerance and cooperation (<xref ref-type="fig" rid="pone.0180754.g004">Fig 4</xref>).</p>
<fig id="pone.0180754.g004" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.g004</object-id>
<label>Fig 4</label>
<caption>
<title>A set of symmetric equilibria of the Markov game on the plane tolerance-betrayal, reciprocal cooperation (theoretical result).</title>
<p>Reciprocal cooperation γ (y-axis) as a function of the tolerance to the defection α(x-axis) for every experiment.</p>
</caption>
<graphic mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.g004" xlink:type="simple"/>
</fig>
<p>The upper point (0, 1) in <xref ref-type="fig" rid="pone.0180754.g004">Fig 4</xref> corresponds to the standard tit-for-tat strategy with 100% reciprocal cooperation and zero tolerance to defection [<xref ref-type="bibr" rid="pone.0180754.ref049">49</xref>]. It is not an interior point of the space of strategic parameters, so the stationary distribution for the pairs of such strategies is determined uniquely and depends on the initial conditions. Without going into detail, let us assume that the participants always make a move cooperatively in the first round, then the pair of tit-for-tat strategies leads to complete cooperation.</p>
<p>For us, the equilibria with maximum tolerance to defection (0.3, 0.8) are of particular importance. We will treat a section of the hyperbolic curve below this point as equilibria before Socialization, and above the point as equilibria after Socialization. Let us pay attention to the fact that high levels of reciprocal cooperation (over 80%) are realized only during rather low tolerance.</p>
</sec>
</sec>
<sec id="sec010" sec-type="conclusions">
<title>Results and discussion</title>
<p>Let us apply the theoretical calculations obtained to the experimental data. We will consider the data on the PD before and after Socialization.</p>
<p><xref ref-type="table" rid="pone.0180754.t005">Table 5</xref> displays the cases of cooperative and non-cooperative moves, reciprocal cooperation and tolerance to defection in PD before and after Socialization. There Ncoop is the number of a partner's cooperative moves in the previous round; Nrecoop the number of cooperative moves in response to the cooperation in the previous round; Ndefault the number of a partner's non-cooperative moves in the previous round; and Ntolerant the number of cooperative moves after the partner's defection in the previous round.</p>
<table-wrap id="pone.0180754.t005" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.t005</object-id>
<label>Table 5</label> <caption><title>Number of cooperative and non-cooperative moves, reciprocal cooperation and tolerance to defection in PD before and after Socialization.</title></caption>
<alternatives>
<graphic id="pone.0180754.t005g" mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.t005" xlink:type="simple"/>
<table>
<colgroup>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
</colgroup>
<thead>
<tr>
<th align="center" rowspan="2">Experiment date</th>
<th align="center" colspan="4">Before Socialization</th>
<th align="center" colspan="4">After Socialization</th>
</tr>
<tr>
<th align="center">Ncoop</th>
<th align="center">Nrecoop</th>
<th align="center">Ndefault</th>
<th align="center">Ntolerant</th>
<th align="center">Ncoop</th>
<th align="center">Nrecoop</th>
<th align="center">Ndefault</th>
<th align="center">Ntolerant</th>
</tr>
</thead>
<tbody>
<tr style="border-top:thick">
<td align="center">15.09.2015</td>
<td align="center">31</td>
<td align="center">7</td>
<td align="center">221</td>
<td align="center">23</td>
<td align="center">70</td>
<td align="center">61</td>
<td align="center">26</td>
<td align="center">12</td>
</tr>
<tr>
<td align="center">21.09.2015</td>
<td align="center">19</td>
<td align="center">2</td>
<td align="center">101</td>
<td align="center">14</td>
<td align="center">53</td>
<td align="center">39</td>
<td align="center">43</td>
<td align="center">17</td>
</tr>
<tr>
<td align="center">28.09.2015</td>
<td align="center">50</td>
<td align="center">25</td>
<td align="center">70</td>
<td align="center">23</td>
<td align="center">131</td>
<td align="center">100</td>
<td align="center">73</td>
<td align="center">30</td>
</tr>
<tr>
<td align="center">05.10.2015</td>
<td align="center">28</td>
<td align="center">4</td>
<td align="center">92</td>
<td align="center">21</td>
<td align="center">119</td>
<td align="center">90</td>
<td align="center">85</td>
<td align="center">28</td>
</tr>
<tr>
<td align="center">09.10.2015</td>
<td align="center">43</td>
<td align="center">18</td>
<td align="center">77</td>
<td align="center">22</td>
<td align="center">109</td>
<td align="center">73</td>
<td align="center">95</td>
<td align="center">34</td>
</tr>
<tr>
<td align="center">12.10.2015</td>
<td align="center">52</td>
<td align="center">25</td>
<td align="center">68</td>
<td align="center">25</td>
<td align="center">147</td>
<td align="center">120</td>
<td align="center">57</td>
<td align="center">25</td>
</tr>
<tr>
<td align="center">19.10.2015</td>
<td align="center">43</td>
<td align="center">20</td>
<td align="center">77</td>
<td align="center">20</td>
<td align="center">76</td>
<td align="center">65</td>
<td align="center">20</td>
<td align="center">13</td>
</tr>
<tr>
<td align="center">26.10.2015</td>
<td align="center">26</td>
<td align="center">9</td>
<td align="center">94</td>
<td align="center">15</td>
<td align="center">112</td>
<td align="center">91</td>
<td align="center">92</td>
<td align="center">22</td>
</tr>
</tbody>
</table>
</alternatives>
<table-wrap-foot>
<fn id="t005fn001"><p>The table contains the number of a partner's cooperative moves in the previous round (Ncoop); the number of cooperative moves in response to the cooperation in the previous round (Nrecoop); the number of a partner's non-cooperative moves in the previous round (Ndefault); and the number of cooperative moves after the partner's defection in the previous round (Ntolerant) for all experiments before and after Socialization.</p></fn>
</table-wrap-foot>
</table-wrap>
<p>This data allow us to assess the parameters α and γ for each experiment before and after Socialization. It is natural to assess value α as <inline-formula id="pone.0180754.e013"><alternatives><graphic id="pone.0180754.e013g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e013" xlink:type="simple"/><mml:math display="inline" id="M13"><mml:mrow><mml:mfrac><mml:mrow><mml:mi mathvariant="normal">Ntolerant</mml:mi></mml:mrow><mml:mrow><mml:mi mathvariant="normal">Ndefault</mml:mi></mml:mrow></mml:mfrac></mml:mrow></mml:math></alternatives></inline-formula>, and value γ as <inline-formula id="pone.0180754.e014"><alternatives><graphic id="pone.0180754.e014g" mimetype="image" position="anchor" xlink:href="info:doi/10.1371/journal.pone.0180754.e014" xlink:type="simple"/><mml:math display="inline" id="M14"><mml:mrow><mml:mfrac><mml:mrow><mml:mi mathvariant="normal">Nrecoop</mml:mi></mml:mrow><mml:mrow><mml:mi mathvariant="normal">Ncoop</mml:mi></mml:mrow></mml:mfrac></mml:mrow></mml:math></alternatives></inline-formula>. This assessment is presented in <xref ref-type="table" rid="pone.0180754.t006">Table 6</xref>.</p>
<table-wrap id="pone.0180754.t006" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.t006</object-id>
<label>Table 6</label> <caption><title>Assessment of the parameters of reciprocal Socialization and tolerance to defection according to the results of the experiments before and after Socialization.</title></caption>
<alternatives>
<graphic id="pone.0180754.t006g" mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.t006" xlink:type="simple"/>
<table>
<colgroup>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
</colgroup>
<thead>
<tr>
<th align="left" rowspan="2" style="background-color:#FFFFFF">Experiment date</th>
<th align="center" colspan="2" style="background-color:#FFFFFF">Before Socialization</th>
<th align="center" colspan="2" style="background-color:#FFFFFF">After Socialization</th>
</tr>
<tr>
<th align="center" style="background-color:#FFFFFF">Alpha, α</th>
<th align="center" style="background-color:#FFFFFF">Gamma, γ</th>
<th align="center" style="background-color:#FFFFFF">Alpha, α</th>
<th align="center" style="background-color:#FFFFFF">Gamma, γ</th>
</tr>
</thead>
<tbody>
<tr style="border-top:thick">
<td align="center" style="background-color:#FFFFFF">15.09.2015</td>
<td align="center" style="background-color:#FFFFFF">0.104</td>
<td align="center" style="background-color:#FFFFFF">0.226</td>
<td align="center" style="background-color:#FFFFFF">0.462</td>
<td align="center" style="background-color:#FFFFFF">0.871</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">21.09.2015</td>
<td align="center" style="background-color:#FFFFFF">0.139</td>
<td align="center" style="background-color:#FFFFFF">0.105</td>
<td align="center" style="background-color:#FFFFFF">0.395</td>
<td align="center" style="background-color:#FFFFFF">0.736</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">28.09.2015</td>
<td align="center" style="background-color:#FFFFFF">0.329</td>
<td align="center" style="background-color:#FFFFFF">0.500</td>
<td align="center" style="background-color:#FFFFFF">0.411</td>
<td align="center" style="background-color:#FFFFFF">0.763</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">05.10.2015</td>
<td align="center" style="background-color:#FFFFFF">0.228</td>
<td align="center" style="background-color:#FFFFFF">0.143</td>
<td align="center" style="background-color:#FFFFFF">0.329</td>
<td align="center" style="background-color:#FFFFFF">0.756</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">09.10.2015</td>
<td align="center" style="background-color:#FFFFFF">0.286</td>
<td align="center" style="background-color:#FFFFFF">0.419</td>
<td align="center" style="background-color:#FFFFFF">0.358</td>
<td align="center" style="background-color:#FFFFFF">0.670</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">12.10.2015</td>
<td align="center" style="background-color:#FFFFFF">0.368</td>
<td align="center" style="background-color:#FFFFFF">0.481</td>
<td align="center" style="background-color:#FFFFFF">0.439</td>
<td align="center" style="background-color:#FFFFFF">0.816</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">19.10.2015</td>
<td align="center" style="background-color:#FFFFFF">0.260</td>
<td align="center" style="background-color:#FFFFFF">0.465</td>
<td align="center" style="background-color:#FFFFFF">0.650</td>
<td align="center" style="background-color:#FFFFFF">0.855</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">26.10.2015</td>
<td align="center" style="background-color:#FFFFFF">0.160</td>
<td align="center" style="background-color:#FFFFFF">0.346</td>
<td align="center" style="background-color:#FFFFFF">0.239</td>
<td align="center" style="background-color:#FFFFFF">0.813</td>
</tr>
</tbody>
</table>
</alternatives>
<table-wrap-foot>
<fn id="t006fn001"><p>The table contains tolerance to defection (Alpha) and reciprocal cooperation (Gamma) for all experiments before and after Socialization.</p></fn>
</table-wrap-foot>
</table-wrap>
<p>Now let us placed the obtained pairs of assessment from <xref ref-type="table" rid="pone.0180754.t006">Table 6</xref> on the plane (α,γ) together with a section of the hyperbolic curve falling within the unit square (<xref ref-type="fig" rid="pone.0180754.g005">Fig 5</xref>).</p>
<fig id="pone.0180754.g005" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.g005</object-id>
<label>Fig 5</label>
<caption>
<title>Tolerance to defection and reciprocal cooperation: Theory and experiment.</title>
<p>Reciprocal cooperation γ (y-axis) as a function of the tolerance of the defection α (x-axis) for every experiments: comparison of the theoretical and experimental data.</p>
</caption>
<graphic mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.g005" xlink:type="simple"/>
</fig>
<p>From the data shown in <xref ref-type="fig" rid="pone.0180754.g005">Fig 5</xref> we can formulate the following results:</p>
<p>Result 1. An increase in the responding cooperation level after Socialization.</p>
<p>The separation of the points in <xref ref-type="fig" rid="pone.0180754.g005">Fig 5</xref> into two vertical clusters is evident.</p>
<p>Result 2. Observable tolerance to betrayal exceeds theoretical tolerance in the equilibrium in nearly all the experiments (15 points out of 16).</p>
<p>All the points except one in <xref ref-type="fig" rid="pone.0180754.g005">Fig 5</xref> lie to the right of the hyperbolic curve.</p>
<p>Result 3. More than a third of the experiments (6 points out of 16) are consistent with the theory.</p>
<p>Result 4. Almost all the experimental data are in a position to the ε-equilibria for the iterated PD in Markov strategies.</p>
<p>Let us set off the horizontal distance (tolerance) from the experimental points to the hyperbolic curve to prove results 3 and 4.</p>
<p>The experiments in which the distance to the theoretical equilibria of tolerance is less than 0.1 are highlighted in <xref ref-type="table" rid="pone.0180754.t007">Table 7</xref>.</p>
<table-wrap id="pone.0180754.t007" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.t007</object-id>
<label>Table 7</label> <caption><title>The distance of tolerance to defection from the theoretical equilibria to the experimental data before and after Socialization.</title></caption>
<alternatives>
<graphic id="pone.0180754.t007g" mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.t007" xlink:type="simple"/>
<table>
<colgroup>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
</colgroup>
<thead>
<tr>
<th align="left">Experiment date</th>
<th align="center">Before Socialization</th>
<th align="center">After Socialization</th>
</tr>
</thead>
<tbody>
<tr style="border-top:thick">
<td align="center">15.09.2015</td>
<td align="center"><underline><bold><italic>0</italic>.<italic>03269024</italic></bold></underline></td>
<td align="center">0.185075</td>
</tr>
<tr>
<td align="center">21.09.2015</td>
<td align="center">0.14237892</td>
<td align="center">0.106521</td>
</tr>
<tr>
<td align="center">28.09.2015</td>
<td align="center">0.11205629</td>
<td align="center">0.119693</td>
</tr>
<tr>
<td align="center">05.10.2015</td>
<td align="center">0.20802336</td>
<td align="center"><underline><bold><italic>0</italic>.<italic>038593</italic></bold></underline></td>
</tr>
<tr>
<td align="center">09.10.2015</td>
<td align="center">0.10754084</td>
<td align="center"><underline><bold><italic>0</italic>.<italic>081434</italic></bold></underline></td>
</tr>
<tr>
<td align="center">12.10.2015</td>
<td align="center">0.1597411</td>
<td align="center">0.148679</td>
</tr>
<tr>
<td align="center">19.10.2015</td>
<td align="center"><underline><bold><italic>0</italic>.<italic>05905651</italic></bold></underline></td>
<td align="center">0.367935</td>
</tr>
<tr>
<td align="center">26.10.2015</td>
<td align="center"><underline><bold><italic>0</italic>.<italic>01923364</italic></bold></underline></td>
<td align="center"><underline><bold><italic>-0</italic>.<italic>0512</italic></bold></underline></td>
</tr>
</tbody>
</table>
</alternatives>
<table-wrap-foot>
<fn id="t007fn001"><p>The table displays the distance of tolerance to defection from the theoretical equilibria to the experimental data before and after Socialization.</p></fn>
</table-wrap-foot>
</table-wrap>
<p>Now let us check how significant the deviation in profit is from the theoretical equilibria. To do this, we will make the following calculation.</p>
<list list-type="order">
<list-item><p>Let us calculate the point on the hyperbolic curve with the same reciprocal cooperation for each experimental point.</p></list-item>
<list-item><p>For each point on the hyperbolic curve let us calculate the values of equilibrium of the player's profit.</p></list-item>
<list-item><p>Let us treat each experimental point as a deviation of one of the players on tolerance to betrayal from the equilibrium, considering that the other player adheres to the equilibrium. Let us calculate the decrease in the profit of the deviated player in the percentage of his equilibrium profit.</p></list-item>
</list>
<p>The results of this calculation are presented in <xref ref-type="table" rid="pone.0180754.t008">Table 8</xref>.</p>
<table-wrap id="pone.0180754.t008" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.t008</object-id>
<label>Table 8</label> <caption><title>The deviation of the winning from the theoretical equilibria to the experimental data before and after Socialization.</title></caption>
<alternatives>
<graphic id="pone.0180754.t008g" mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.t008" xlink:type="simple"/>
<table>
<colgroup>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
<col align="left" valign="middle"/>
</colgroup>
<thead>
<tr>
<th align="center" style="background-color:#FFFFFF">Experiment date</th>
<th align="center" style="background-color:#FFFFFF">Before Socialization</th>
<th align="center" style="background-color:#FFFFFF">After Socialization</th>
</tr>
</thead>
<tbody>
<tr style="border-top:thick">
<td align="center" style="background-color:#FFFFFF">15.09.2015</td>
<td align="center" style="background-color:#FFFFFF">0.03%</td>
<td align="center" style="background-color:#FFFFFF">0.31%</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">21.09.2015</td>
<td align="center" style="background-color:#FFFFFF">0.92%</td>
<td align="center" style="background-color:#FFFFFF">0.16%</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">28.09.2015</td>
<td align="center" style="background-color:#FFFFFF">0.25%</td>
<td align="center" style="background-color:#FFFFFF">0.19%</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">05.10.2015</td>
<td align="center" style="background-color:#FFFFFF">1.68%</td>
<td align="center" style="background-color:#FFFFFF">0.02%</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">09.10.2015</td>
<td align="center" style="background-color:#FFFFFF">0.26%</td>
<td align="center" style="background-color:#FFFFFF">0.11%</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">12.10.2015</td>
<td align="center" style="background-color:#FFFFFF">0.51%</td>
<td align="center" style="background-color:#FFFFFF">0.25%</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">19.10.2015</td>
<td align="center" style="background-color:#FFFFFF">0.08%</td>
<td align="center" style="background-color:#FFFFFF">1.01%</td>
</tr>
<tr>
<td align="center" style="background-color:#FFFFFF">26.10.2015</td>
<td align="center" style="background-color:#FFFFFF">0.01%</td>
<td align="center" style="background-color:#FFFFFF">0.04%</td>
</tr>
</tbody>
</table>
</alternatives>
<table-wrap-foot>
<fn id="t008fn001"><p>The table shows the deviation of the winning from the theoretical equilibria to the experimental data before and after Socialization.</p></fn>
</table-wrap-foot>
</table-wrap>
<p>For clarity, let us order the values obtained in <xref ref-type="table" rid="pone.0180754.t008">Table 8</xref> in descending order and present them graphically (<xref ref-type="fig" rid="pone.0180754.g006">Fig 6</xref>).</p>
<fig id="pone.0180754.g006" position="float">
<object-id pub-id-type="doi">10.1371/journal.pone.0180754.g006</object-id>
<label>Fig 6</label>
<caption>
<title>Deviation of experiments from the equilibria on winning in percentages.</title>
<p>The deviation in winning in the experimental data from the theoretical equilibrium (y-axis) during the experiments in decreasing order of the deviations from the equilibria in winning (x-axis).</p>
</caption>
<graphic mimetype="image" position="float" xlink:href="info:doi/10.1371/journal.pone.0180754.g006" xlink:type="simple"/>
</fig>
<p><xref ref-type="fig" rid="pone.0180754.g006">Fig 6</xref> shows that the maximum deviation from the equilibria on winning is only 1.7%, and this deviation is no more than 0.5% in 75% of cases. This means that nearly all the experiments are in a position of ε-equilibria for the iterated PD in Markov strategies. This result is fundamental.</p>
</sec>
<sec id="sec011" sec-type="conclusions">
<title>Conclusions</title>
<p>This research applied a QRE model to the results of an experiment designed to study cooperation and trust under the influence of social interaction. The resulting data were divided into two categories: before and after social interaction. The peculiarity of the results is that the data on cooperation after Socialization in the Socialization stage are significantly different from those of the experiments in PD. The calculations showed that the behavior of the participants before Socialization could be described with the QRE concept, which is an accepted deviation from the concept of Nash equilibrium that weakens the requirements of the best response. However, the standard QRE approach cannot be applied to describing the behavior of the participants after Socialization. Therefore, we have proposed a variant of the description of equilibria in the iterated PD in Markov strategies. For this game repeated in Markov strategies, we managed to explicitly find all the equilibria with a positive probability of reciprocal cooperation and tolerance to betrayal. The primary result is that all the experiments are in a position of ε-equilibria for the repeated PD game in Markov strategies. There remain questions of theoretical justification of the results of such games as the Trust Game and Ultimatum Game, the experimental data of which do not correspond to known theoretical game models in the framework of our research on the influence of social interaction.</p>
</sec>
</body>
<back>
<ack>
<p>We thank Rinat Yaminov for writing the programing code for experiments, Anna Sedush and Alexander Chaban for technical help in conducting experiments at Tomsk State University, Olga Menshikova for thoughtful comments and advices. This research was supported by The Tomsk State University competitiveness improvement program.</p>
</ack>
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